changed fitness impact

model/FDM.nlogo

@@ -135,7 +135,7 @@ to setup
 
   set no-output temp-no-output
 
-  set model-version "V1.3.1"
+  set model-version "V1.4.0"
 
   if behaviorspace-run-number > 0 [ random-seed behaviorspace-run-number ]
 

model/flies.nls

@@ -565,24 +565,35 @@ to fly-activities
   let fertile-female-adult-flies female-adult-flies with [ partner-search = TRUE ]
 
   ask fertile-female-adult-flies [
-
-    let fertile-male-adult-flies-here flies-here with [ sex = "m" and mode = "adult" and partner-search = TRUE ]
+
+    let mating-chance random-float 1
+    let fertile-male-adult-flies-here flies-here with [ sex = "m" and mode = "adult" and partner-search = TRUE and fitness >= mating-chance ]
 
     ; if female fly detects male fly on same patch => start fertilization
     if any? fertile-male-adult-flies-here [
-      set partner-search FALSE
-      set fertilization-tick ticks
-      set eggs eggs-per-cycle
+
       let partner n-of 1 fertile-male-adult-flies-here 
-      ; "[genotype] of partner" and "[fitness] of partner" return lists
-      set genotype-of-partner first [genotype] of partner
-      set fitness-of-partner first [fitness] of partner
-      set combined-fitness ( fitness * fitness-of-partner )
-      set color orange
       ask partner [
         set partner-search FALSE
         set fertilization-tick ticks 
       ]
+
+      let mating-success random-float 1
+      ifelse ( fitness >= mating-success ) [
+
+        set partner-search FALSE
+        set fertilization-tick ticks
+        set eggs eggs-per-cycle
+        ; "[genotype] of partner" and "[fitness] of partner" return lists
+        set genotype-of-partner first [genotype] of partner
+        set fitness-of-partner first [fitness] of partner
+        set combined-fitness ( fitness * fitness-of-partner )
+        set color orange
+
+      ] [
+        start-immature-state
+      ]
+
     ]     
 
   ]
@@ -620,7 +631,8 @@ to fly-activities
         ]
 
         ; fitness rate
-        if random-float 1 > combined-fitness [ set lay-egg FALSE ]
+        ; deactivated. new impact of fitness: mating chance(m)/success(f)
+        ; if random-float 1 > combined-fitness [ set lay-egg FALSE ]
 
         ifelse lay-egg [
 

model/verification.nls

@@ -1736,45 +1736,90 @@ to-report test-fitness [ verification-file ]
   let log-header "test-fitness: "
   let log-content ""
 
-  ; check if cherry and fruit number get reduced if egg gets not layed due to fitness
-
-  refresh-world
-  ; grow cherries
-  ask trees [
-    set grown-cherries 5
-    set occupied-cherries 0
+  ; FIRST TEST 
+  ; 300 trials
+  ; 1 female / 1 male fly
+  ; female fitness 1
+  ; male fitness 0.4
+  ; mating success should be around 40%
+
+  let trials 300
+  let trial 1
+  let matings 0
+  while [ trial <= trials ] [
+
+    refresh-world
+    random-seed new-seed
+
+    set init-pop 2
+    start-season
+    ask flies with-min [who] [
+      set sex "f"
+      set fitness 1
+      move-to min-one-of trees [who] ; move to tree with lowest id
+    ]
+    ask flies with-max [who] [
+      set sex "m"
+      set fitness 0.4
+      move-to min-one-of trees [who] ; move to tree with lowest id
+    ]
+
+    fly-activities
+
+    ; check mating
+    if count flies with [ sex = "f" and eggs > 0 ] > 0 [
+      set matings ( matings + 1 ) 
+    ]
+
+    set trial ( trial + 1 ) 
   ]
-  set cherries-available TRUE
-  ; grow wildberries
-  ask wildberry-plants [
-    set grown-wildberries 5
-    set occupied-wildberries 0
+  ; Most of the time the result should be between .38 and .42, but due to low total trials it may happen that the deviation is slightly higher. Therefore, .36 and .44 chosen for boundaries.
+  if ( matings / trials ) < 0.36 or ( matings / trials ) > 0.44 [
+     set log-content ( word log-content "CHECK FITNESS: first test failed - mating success not around 40% (" round( matings / trials * 100 ) "%); " )
   ]
-  set wildberries-available TRUE
-  ; create flies
-  set init-pop 1
-  start-season
-  ; modify flies
-  ask flies [
-    set sex "f"
-    set ready-to-lay-egg TRUE
-    set eggs 1
-    set genotype "++"
-    set genotype-of-partner "++"
-    set fitness 0
-    set fitness-of-partner 0
-    set combined-fitness 0
-    set fertilization-tick 0
-    move-to min-one-of trees [who] ; move to tree with lowest id
+
+  ; SECOND TEST 
+  ; 300 trials
+  ; 1 female / 1 male fly
+  ; female fitness 0.6
+  ; male fitness 1
+  ; mating success should be around 60%
+
+  set trials 300
+  set trial 1
+  set matings 0
+  while [ trial <= trials ] [
+
+    refresh-world
+    random-seed new-seed
+
+    set init-pop 2
+    start-season
+    ask flies with-min [who] [
+      set sex "f"
+      set fitness 0.6
+      move-to min-one-of trees [who] ; move to tree with lowest id
+    ]
+    ask flies with-max [who] [
+      set sex "m"
+      set fitness 1
+      move-to min-one-of trees [who] ; move to tree with lowest id
+    ]
+
+    fly-activities
+
+    ; check mating
+    if count flies with [ sex = "f" and eggs > 0 ] > 0 [
+      set matings ( matings + 1 ) 
+    ]
+
+    set trial ( trial + 1 ) 
   ]
-  fly-activities
-  ; check values
-  if count flies with [ mode = "adult" ] != 1 [
-    set log-content word log-content "CHECK FITNESS: wrong number of adult flies; "
+  ; Most of the time the result should be between .56 and .64, but due to low total trials it may happen that the deviation is slightly higher. Therefore, .56 and .64 chosen for boundaries.
+  if ( matings / trials ) < 0.56 or ( matings / trials ) > 0.64 [
+     set log-content ( word log-content "CHECK FITNESS: second test failed - mating success not around 60% (" round( matings / trials * 100 ) "%); " )
   ]
-  if count flies with [ mode = "stationary" ] != 0 [
-    set log-content word log-content "CHECK FITNESS: wrong number of stationary flies; "
-  ]    
+
 
   write-verification-result-to-file verification-file log-header log-content
   ifelse length log-content > 0 [ report FALSE ] [ report TRUE ]